CNN  — 

Fierce fighting across Sudan has left hopes for a peaceful transition to civilian rule in tatters. Forces loyal to two rival generals are vying for control, and as is so often the case, civilians have suffered the most.

At least 459 people have been killed and more than 4,000 injured in the unrest so far, according to the World Health Organization, while parts of the capital Khartoum have become a war zone.

A 72-hour ceasefire agreed on by Sudan’s Armed Forces (SAF) and Sudan’s Rapid Support Forces (RSF) came into effect at midnight local time on Tuesday (Monday 6 p.m. ET).

Volker Perthes, the UN Secretary General’s Special Representative for Sudan, said Tuesday that the ceasefire “seems to be holding in some parts; however, reports of sporadic shooting are still coming in as well as reports of relocation of troops.”

Doctors’ organizations and several eyewitnesses said medical facilities were being bombarded with military strikes in targeted attacks. Both sides later denied shelling hospitals, in comments to CNN.

The unrest entered its 12th day on Wednesday as efforts to evacuate foreign diplomats and citizens intensified.

Here’s what you need to know.

A power struggle is at the center of the fighting

At the heart of the clashes are two men: Sudan’s military ruler and head of the army Abdel Fattah al-Burhan and General Mohamed Hamdan Dagalo (widely known as Hemedti), the country’s deputy and head of the Rapid Support Forces (RSF) paramilitary group.

Until recently, the two men were allies who worked together to topple ousted Sudanese President Omar al-Bashir in 2019 and played a pivotal role in the military coup in 2021.

However, tensions arose during negotiations to integrate the RSF into the country’s military as part of plans to restore civilian rule.

The key question: who would be subordinate to whom under the new hierarchy.

These hostilities, sources told CNN, are the culmination of what both parties view as an existential fight for dominance.

Smoke is seen over Khartoum on April 22, 2023 after days of fighting in the Sudanese capital.

Sudan is no stranger to upheaval

It is difficult to overstate how seismic Bashir’s overthrow was. He had led the country for nearly three decades when popular protests that began over soaring bread prices toppled him from power.

During his rule, South Sudan split from the north while the International Criminal Court issued an arrest warrant for Bashir alleged war crimes in Darfur, a separatist Western region.

After Bashir’s ouster, Sudan was ruled by an uneasy alliance between the military and civilian groups.

That all ended in 2021, when the power-sharing government was dissolved by armed forces.

Former Sudanese minister Ahmed Haroun, wanted by the International Criminal Court (ICC) for war crimes, said he and a number of former regime figures, left prison in Khartoum after chaos hit the facility on Sunday. Unconfirmed reports on Wednesday claimed Bashir was among those released.

However, the media office of Sudan’s Police and sources familiar with the matter told CNN Bashir remains in the custody of the Sudan Armed Forces at a military hospital in Omdurman, west of Khartoum.

Passengers fleeing war-torn Sudan disembark at a bus station near the Egyptian city of Aswan, on April 25, 2023.

The RSF has a controversial past

The Rapid Support Forces are the preeminent paramilitary group in Sudan, whose leader, Dagalo, has enjoyed a rapid rise to power.

During Sudan’s Darfur conflict in the early 2000s, he was the leader of Sudan’s notorious Janjaweed forces, implicated in human rights violations and atrocities.

An international outcry saw Bashir formalize the group into paramilitary forces known as the Border Intelligence Units.

In 2007, its troops became part of the country’s intelligence services and, in 2013, Bashir created the RSF, a paramilitary group overseen by him and led by Dagalo.

Dagalo turned against Bashir in 2019, but not before his forces opened fire on an anti-Bashir, pro-democracy sit-in in Khartoum, killing at least 118 people.

He was later appointed deputy of the transitional Sovereign Council that ruled Sudan in partnership with civilian leadership.

Russian involvement

Russia’s mercenary group, Wagner, has involved itself in the conflict by boosting the RSF’s missile supplies, Sudanese and regional diplomatic sources told CNN.

The sources said the surface-to-air missiles have significantly buttressed RSF paramilitary fighters and their leader Dagalo.

In bordering Libya, where a Wagner-backed rogue general, Khalifa Haftar, controls swathes of land, satellite imagery supports these claims, showing an unusual uptick in activity on Wagner bases.

The bolster in support comes amid deepening ties between Moscow and Sudan’s military leadership.

A July 2022 CNN investigation found that Sudan’s military leaders granted Russia access to the east African country’s gold riches in exchange for military and political support.

Dagalo’s forces were a key recipient of Russian training and weaponry, and Sudan’s military leader Burhan is also believed by CNN’s Sudanese sources to have been backed by Russia, before international pressure forced him to publicly disavow the presence of the Russian mercenary group Wagner, in Sudan.

The two rivals mirror each other

Burhan is essentially Sudan’s leader. At the time of Bashir’s toppling, Burhan was the army’s inspector general.

His career has run an almost parallel course to Dagalo’s.

He also rose to prominence in the 2000’s for his role in the dark days of the Darfur conflict, where the two men are believed to have first came into contact.

Al-Burhan and Dagalo both cemented their rise to power by currying favor with the Gulf powerhouses.

They commanded separate battalions of Sudanese forces, who were sent to serve with the Saudi-led coalition forces in Yemen.

Now they find themselves locked in a power struggle.

‘Attempted coup’

Speaking with CNN, Burhan, characterized the RSF offensive as an “attempted coup.”

“This is an attempted coup and rebellion against the state,” Burhan told CNN by phone. He said that RSF leader Dagalo had “mutinied” against the state, and if captured, would be tried in court of law.

Burhan alleged in the CNN interview that the RSF had tried to “capture me and kill me.”

When asked about that claim, an RSF spokesperson told CNN that the group was “seeking to capture him” and bring him to justice for “many treasons acts against the Sudanese people.”

“We are fighting for all Sudanese people,” the spokesperson said in a statement. “We [are] going to bring all responsible parties to justice and give them a fair trial.”

When challenged on why the Sudanese people should trust him, given his former alliance with Dagalo, Burhan told CNN: “The Sudanese Army is the people’s army.

“It is not owned by specific people or specific organizations,” he said. “It is a national institution, which is tasked with defending Sudan.”

Sudan faces an uncertain future

Where the fighting will end is unclear. Both sides claim control over key sites and fighting has been reported across the country in places far from the capital Khartoum.

While various official and non-official estimates place the Sudanese armed forces at around 210-220,000, the RSF are believed to number approximately 70,000 but are better trained and better equipped.

International powers have expressed alarm, with the UN Security Council holding its first meeting on the crisis in Sudan on Monday. Apart from concerns over civilians there are likely other motivations at play – Sudan is resource-rich and strategically located.

Sudan’s neighbors Egypt and South Sudan have offered to mediate, but in the meantime all that is certain is more misery for the Sudanese people.

CNN’s AnneClaire Stapleton contributed reporting.