For a moment, the surrender took a dramatic turn when the last holdout refused to follow the other three and continued the standoff at the Malheur National Wildlife Refuge's headquarters in Harney County.
Fry said, "I am walking to them right now" as the other people on the phone call said, "Hallelujah, keep walking."
Through the open phone line, listeners could hear officials as they arrested the man, saying "nice and easy, hands behind your back."
The end of the siege came after the armed occupiers earlier indicated they would turn themselves in. By morning's end, they fulfilled that pledge.
Moments before Fry surrendered, he said he would walk out of the refuge -- if everyone exclaimed "Hallelujah!" said Harney County Sheriff David Ward.
Just about everyone did, even SWAT team members, the sheriff said.
Last month, one of the leading occupiers was killed by authorities in a highway incident.
Before the final surrender, federal agents arrested the figurehead of the protesters' movement: Nevada rancher Cliven Bundy, who didn't participate in the occupation but is the father of arrested standoff leader Ammon Bundy.
For now, the arrests of the Bundys and the end of occupation put the dispute in the hands of the courts. The Bundys and their supporters argue that the federal government has exceeded the boundaries of the Constitution.
Cliven Bundy announced on Wednesday he was heading to Oregon, according to a Facebook page for his Nevada ranch.
In 2014, he came to the national spotlight in a showdown with the federal Bureau of Land Management over grazing rights for his cattle.
"It's time!" the Facebook post said. "Cliven Bundy is headed to the Harney County Resource Center in Burns Oregon."
But Bundy was taken into federal custody in Portland after landing early Thursday, the FBI said.
Bundy to appear in court Thursday
Bundy was charged with six counts relating to the 2014 standoff with the federal government
, according to a criminal complaint filed in U.S. District Court in Nevada.
He was charged with conspiracy to commit an offense against the United States, assault on a federal law officer by use of a deadly and dangerous weapon, interference with commerce by extortion, and obstruction of justice. He was also charged with two counts of use and carry of a firearm in relation to a crime of violence, the complaint said.
He appeared in federal court in Portland and asked for a court-appointed attorney, The Oregonian reported.
The Bundy ranch's Facebook page posted a statement
that Bundy was arrested for charges "related to his standoff with the U.S. Bureau of Land Management in 2014." The post continued, "The charges include a conspiracy charge to interfere with a federal officer ... and a weapons charge."
Bundy's son, Ammon
, was arrested last month.
Long list of alleged occupiers
Authorities identified the last four occupiers arrested Thursday: Fry, 27, of Blanchester, Ohio; Jeff Wayne Banta, 46, of Yerington, Nevada; and Sean Larry Anderson, 47, and Sandra Lynn Anderson, 48, both of Riggins, Idaho, the FBI said.
They are scheduled to be arraigned Friday in federal court in Portland.
No one was injured, and no shots were fired in the last arrests, the FBI said.
During the 41-day occupation, there were as many as "dozens of highly armed militants occupying, visiting and supplying the refuge," the FBI said.
On February 3, a federal grand jury indicted 16 people, including the four who surrendered, the FBI said.
The other 12 people indicted are Dylan Wade Anderson, 34, of Provo, Utah; Brian Cavalier, 44, of Bunkerville, Nevada; Shawna Cox, 59, Kanab, Utah; Duane Leo Ehmer, 45, of Irrigon, Oregon; Kenneth Medenbach, 62, of Crescent, Oregon; Joseph Donald O'Shaughnessy, 45, of Cottonwood, Arizona; Jason S. Patrick, 43, of Bonaire, Georgia; Ryan Waylen Payne, 32, of Anaconda, Montana; Jon Eric Ritzheimer, 32, Peoria, Arizona; and Peter Santilli, 50, of Cincinnati, Ohio, according to the FBI.
Also indicted are Ammon Edward Bundy, 40, of Emmett, Idaho, and his brother Ryan C. Bundy, 43, of Bunkerville, Nevada, the FBI said.
Each are charged with a federal felony count of conspiracy to impede officers of the United States from discharging their official duties through the use of force, intimidation, or threats, the FBI said.
'Continued presence of militia groups'
The impact of the occupation is still playing out in Oregon, authorities said.
In fact, local authorities report the arrival of "militia groups" to Harney County.
Law officers "will now be able to begin the process of clearing booby traps and processing the crime scene," Harney County Court said in a statement.
"In the meantime, the occupation and continued presence of militia groups in Harney County continues to cause division in the county and every community within. Calls for thousands of people to descend on Burns are very troubling," the court statement said Thursday.
Federal authorities called for a return to normalcy, but acknowledged more work ahead.
"The occupation of the Malheur Wildlife Refuge has been a long and traumatic episode for the citizens of Harney County and the members of the Burns Paiute tribe. It is a time for healing, reconciliation amongst neighbors and friends, and allowing for life to get back to normal," U.S. Attorney Billy J. Williams said in a statement.
"Much work is left to assess the crime scene and damage to the refuge and tribal artifacts. We are committed to seeing the job done and to pursue justice for the crimes committed during the illegal occupation," he added.
The U.S. Fish and Wildlife Service and the tribe will assess any damage to the tribe's burial grounds or artifacts dating back thousands of years, said Gregory Bretzing, special agent in charge of FBI's Portland office.
"Over the course of the last month, the people of Harney County have lived through an experience that is both highly emotional and physically exhausting," he said in a statement. "We have seen some residents leave their homes, fearing violence against their families. We have seen the confusion, concern and trouble that the occupiers' actions have caused for this community."
Added Harney County Judge Steven Grasty at a press conference: "We look forward to the day when our community is ours once again."
Sheriff Ward described his community as "torn apart" over the occupation.
"There's been a lot of hurt, there's been been lot of things said, but I don't think there's anything that's been done that can't be worked through," the sheriff said.
News from a conference call
During a purported live stream of a conference call Wednesday night between protesters, activists and conservative Nevada lawmaker Michele Fiore, the occupiers said they were prepared to leave.
The audio was live-streamed on YouTube.
Fiore told those on the call that Mike Arnold -- Ammon Bundy's lawyer, who Fiore said was in the car with her -- spoke with the FBI. She said the agency promised it would stand down Wednesday night and allow her to be at an FBI checkpoint when the occupiers surrendered.
Fiore was indeed present at the surrender, as was Franklin Graham, the nationally known evangelist and son of legendary preacher Billy Graham, authorities said. Graham said he's been talking with "the last four holdouts ... every day by phone for the last week at their request and at the request of the FBI."
The live stream started after the FBI on Wednesday surrounded those occupying the refuge.
According to the agency, one of the remaining occupiers rode outside barricades at the Malheur National Wildlife Refuge. When agents tried to approach him, he sped back to the refuge.
After that, the FBI said agents "moved to contain the remaining occupiers by placing agents at barricades both immediately ahead of and behind the area where the occupiers are camping."
The FBI said no shots were fired.
"The FBI has negotiated with patience and restraint in an effort to resolve the situation peacefully," Bretzing of the FBI said.
"However, we reached a point where it became necessary to take action in a way that best ensured the safety of those on the refuge, the law enforcement officers who are on scene, and the people of Harney County who live and work in this area."
'God has put us on this path'
Earlier on the call, the occupiers seemed concerned that the FBI planned to move in Wednesday night and that it would lead to their deaths. At times, they seemed to fatalistically embrace that outcome.
When one woman -- presumed to be Fiore -- asked two of them about their families, a man responded, "God has put us on this path. Our families are already taken care of; they weren't in our lives much before all this because God made sure we didn't have that to weigh us down so that we could do this."
The people on the phone could be heard debating conditions for which they'd be willing to leave the refuge.
Protest led to armed occupation
Ammon Bundy and others started out demonstrating against the sentencing of Dwight Hammond and his son Steven, ranchers who were convicted of arson on federal lands in Oregon.
But a January 2 march supporting the Hammonds led to the armed occupation of the refuge building, with protesters decrying what they call government overreach when it comes to federal lands.
Bundy and other members of his group were arrested during an incident along a highway last month.
Those law officers weren't wearing any body cameras, the FBI's Bretzing said.